Written by
C. David Kreger
Introduction
Homo ergaster is one of the more problematic of
somewhat accepted species designations currently tossed
around in anthropological literature. Each individual
researcher that sees ergaster as a valid taxon
sees different specimens as belonging or not belonging
to the taxon. Many researchers deny any validity to the
species at all. On the whole though, most researchers
see too little difference between ergaster and
erectus to form the basis of a species of the former,
separated from the latter. As a general rule of thumb,
one can consider most attributed ergaster specimens
to be early erectus geographically confined to
Africa (however, this is not a hard and fast rule).
The
taxon ergaster was first described in 1975 by
C. Groves and V. Mazak. The specimen attributed as the
type specimen was ER 992, an isolated mandible. Since
then, other specimens have been attributed by various
authors to ergaster, with most researchers placing
the same fossils in erectus. Those who see it
as a valid taxon tend to see it as more closely resembling
modern H. sapiens than does H. erectus.
They tend to see ergaster as a direct ancestor
of modern humans with erectus being an evolutionary
dead-end. Many Out of Africa supporters use this taxon
as evidence that Asian and European specimens did not
contribute genetically to the modern human genome, but
this claim is very weak.
Diagnostic Features
The type specimen for ergaster is KNM-ER 992.
Groves and Mazak claimed that the mandible was significantly
different from H. erectus to deserve its own
species designation. However, they did not compare it
to H. habilis, and the mandible may actually
belong to that taxon. The specimen showed some periodontal
disease, as seen by absorption of bone around the roots
of the teeth. The mandibular symphysis also shows strong
markings for the digastric muscle (important for swallowing
and vocalization), which some people have interpreted
as proof of language by this time.
One
of the most spectacular and important paleoanthropological
finds in recent years was the Nariokotome Boy (KNM-WT
15000), by a team of researchers led by Richard Leakey
and Alan Walker. This find represents the most complete
early hominid ever found, with almost the entire cranium,
and most of the postcranial material intact. This specimen
has been attributed as a male ergaster by some,
though most place it in H. erectus, and that
is where it will be discussed in detail. Other specimens
that have been attributed to ergaster include KNM-ER
3733, SK 847, and KNM-ER 3883.
Several
researchers have tried to define the difference between
ergaster and erectus, P. Andrews and B.
Wood among the more prominent. P. Andrews defined seven
autopomorphies that were characteristic of erectus,
but which ergaster supposedly lacked. However,
G. Bräuer have shown that these are not autopomorphies.
For example, some erectus do not possess these
features, while some ergaster and some habilis
do. Also, some of these autopomorphies are not independent
traits, and should not be considered separately (e.g.,
frontal keel and parietal keel). B. Wood lists seven
traits that link ergaster with H. sapiens,
and that distinguish ergaster from erectus:
- Increased
cranial breadth across the parietal bones.
- Increased
occipital bone length.
- Broader
nasal bones.
- Broader
nasal opening.
- Shorter
cranial base.
- Greater
development of the mandibular symphysis.
- Narrower
M1s and lower canines.
However,
these synapomorphies have been convincingly challenged
by showing them to be present in erectus populations
from Asia. Also, more recent analyses by other researchers
seem to indicate that even if ergaster specimens
are considered as a different taxon than erectus,
the erectus material is still closer to modern
humans cladistically.
Conclusions
In short, H. ergaster does not show significant
promise of lasting as a separate taxon due to several
factors. It has not been shown to be significantly different
from erectus to require the designation of a
new hominid species, and it has not been shown to be
closer to modern humans morphologically as has been
claimed by some. At this time, ergaster basically
means early H. erectus from Africa.
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